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What is another word for backbone?

What's another word for backbone? | Here's a list of synonyms for backbone.

Definition 1: the part of a network that connects other networks together - [noun denoting artifact]

Definition 1: the part of a book's cover that encloses the inner side of the book's pages and that faces outward when the book is shelved - [noun denoting artifact]

Definition 1: fortitude and determination - [noun denoting attribute]

Definition 1: the series of vertebrae forming the axis of the skeleton and protecting the spinal cord - [noun denoting body]

Definition 1: a central cohesive source of support and stability - [noun denoting cognition]